The Compsognathus is a very small (comparatively) theropod dinosaur which lived around 150 million years ago. They were brought to the present on Ibis Island when a faulty Third Energy generator generated too much energy and the only way to stop the reaction was to cause an event which caused a rift in space-time around a large portion of the island. This caused the area from prehistoric times to be swapped with the present area - this brought all past flora and fauna to the present and likewise.
They are generally encountered in large 'swarms' and are surprisingly good at navigating through man-made constructs such as air vents and corridors. While they are not very dangerous and quite passive, they can attack if you get too close or start attacking them first. Usually they are just on the hunt for easy pickings - such as feeding from corpses and other defenceless flora and fauna.
It's usually best to ignore them, but if they must be engaged they will generally only take one hit.
Finally, it's worth mentioning that their size in Dino Crisis isn't far off what they are actually sized, and in fact the Velociraptor in reality was very close to the size of a Compsognathus (in the game, they are vastly different in size). Also, it's now known that Compsognathus were a feathered reptile and thus a close neighbour of a modern bird.
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